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Prior to modern fire prevention measures forest fires were a central distubance factor in boreal forests with a strong impact on forest structure and species assemblages. There has been great variablity in the frequency of fires among different forest types. The occurence of fires has also been affected by the size of the continuous heath forest patch along with several stochastic factors. Dry heath forests may have burned in intervals measured in tens of years while spruce dominated areas have burned only once in hundreds of years. Deciduous herb-rich forest have burned least often.
Fire affects the structure and species assemblage of a forest in many ways. Forest fires boost the creation of deciduous stands and increase the amount of dead wood sometimes in large volumes. Although the amount of dead wood created by prescribed burnings in commercial forests is quite low, many other impacts to microclimate and soil acidity, for example, are alike. Temperature variation in a burned ground layer is wider and the soil is more alkaline than in a non-burned site.
The strongest link between forest fires and species exist for those species for which fire is either a partly or entirely necessary part of their life history. Such a species is, for example, the cranesbill species Geranium bohemicum which seeds require a sudden rise of temperature to above 45 C to germinate. Besides vascular plants, fire dependent species can found among sac and imperfect fungi, possibly even tens of species.
Some 30 insect species may also be found living solely in burned forests. The majority of these are beetles. Many of the insect species feed on the sac and imperfect fungi species.
True fire specialist disappear from a burned site normally within a few years after the fire. Species feeding on dead wood may sustain themselves for longer. The open phase with high volumes of dead wood may last for 20 to 30 years during which saproxylic species are abundant.
Due to scarcity of forest fires and prescribed burnings many fire associated species have become rare. There were altogether 34 extinct and endangered and 11 vulnerable fire species in the last red-list assesment completed in year 2000.
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